Short answer
In mobile first, your styles will inherit upwards from the smallest media query, whereas in desktop first they will inherit down from the largest media query.
With DX8 you can choose whether you want you website to be mobile first or desktop first. This option is within Website settings > Responsive grid settings. Your choice will dictate how the Style builder works.
Read more
Knowledge Base: How do I turn off the breakpoint indicator?
Knowledge Base: What does the breakpoint indicator do?
Knowledge Base: Why doesn't the Responsive grid use Flexbox?
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